Professor II 

Address: 001 Waller Hall, Cook Campus, Rutgers University, New Brunswick, New Jersey

Mailing Address:
Department of Ecology, Evolution & Natural Resources, ENRS Building, Cook College, Rutgers University, 14 College Farm Road, New Brunswick, NJ 08901-8551

  (732) 932-1124    FAX:   (732) 932-8746  E-Mail:

B.S. University of California at Berkeley (Forestry) 1960-1965    

Ph.D. North Carolina State University at Raleigh (Genetics) 1965-1970

Postdoc University of Texas at Austin (Zoology) 1970-1972

Research Interests:
I was originally trained in forestry, but have subsequently worked in anthropology and human genetics, demography and mathematical ecology, bacterial ecology, statistical epidemiology and immunology, and psychometrics. I have worked on quantitative analysis of humans and higher primates, propagule flow in forest trees, taxonomic diversity in forbs and agronomic grasses, clinal variation in insects, ecological niche partitioning in bacteria, and forensic genetics of marine and freshwater fish. Our group is primarily oriented toward mathematical modeling and statistical analysis of theoretical population biology problems, both those in evolution and ecology. Much of our work is theoretical, but computer simulation and data analysis play a role in our program as well. Our concentration on the mathematical aspects does not preclude field work, and we have conducted taxonomic studies of southern pines, genetic anthropology studies of rain forest tribes in Brazil and New Guinea, participated in studies of hydrothermal vents in the Gulf of California, and pollen flow studies in Costa Rica.

We currently work on: (a) the flow of forest tree propagules (pollen and seed) across the landscape, (b) the analysis of spatial organization of genetic variation at the micro-geographic scale (within a single population), (c) the measurement of genetic/species richness/ diversity in evolutionary/ecological practice, and (d) developing statistical and mathematical tools for the newer sorts of molecular genetic data used in systematics. Our students have traditionally defined their own thesis projects, within the broad arena covered by our group.

Selected Publications:
Kaufman SR, Smouse PE and Alvarez-Buylla ER. 1998. Pollen-mediated gene flow and differential male reproductive success in a tropical pioneer tree, Cecropia obtusifolia Bertol. (Moraceae). HEREDITY 81:164-173.

Smouse PE and Peakall R. 1999. Spatial autocorrelation analysis of multi-allele and multi-locus genetic microstructure. HEREDITY 82: 561-573.

Smouse PE, Meagher TR and Kobak CJ. 1999. Parentage analysis in Chamaelirium luteum (L.): Why do some males have disproportionate reproductive contributions? J. EVOL. BIOL. 12:1056-1068.

Smouse PE. 2000. Reticulation inside the species boundary. In: Special Section, Reticulate Evolution, P. Legendre (ed.). J. CLASSIFIC. 17:165-173.

Smouse PE, Dyer RJ, Westfall RD and Sork VL. 2001. Two-generation analysis of pollen flow across a landscape. I. Male gamete heterogeneity among females. EVOLUTION 55:260-271.

Austerlitz F and Smouse PE. 2001. Two-generation analysis of pollen flow across a landscape. II. Relation between fft, pollen dispersal, and inter-female distance. GENETICS 157:851-857.

Austerlitz F and Smouse PE. 2001. Two-generation analysis of pollen flow across a landscape. III. Impact of within-population structure. GENETICAL RESEARCH 78:271-280.

Austerlitz F and Smouse PE. 2002. Two-generation analysis of pollen flow across a landscape. IV. Estimating the dispersal parameter. GENETICS 161:355-363.

Sork VL, Davis F, Smouse PE, Apsit V, Dyer R, Fernandez JM and Kuhn B. 2002. Pollen movement in declining populations of California valley oak, Quercus lobata: Where have all the fathers gone? MOLEC. ECOL. 11:1657-1668.

Irwin AJ, Hamrick JL, Godt MJW and Smouse PE. 2003. A multi-year estimate of the effective pollen donor pool for Albizia julibrissin. HEREDITY 90:187-194.

Stevens MHH, Petchey OL and Smouse PE. 2003. Stochastic relations between species richness and the variability of species composition. OIKOS 103:479-488.

Smouse PE and Sork VL. 2004. Measuring pollen flow in forest trees: A comparison of alternative approaches. FOR .ECOL.  MANAG. 197:21-38.


Current Position:
Professor II, Ecology, Evolution & Natural Resources, Cook College, Rutgers University

UC-Berkeley, BS (Forestry) 65; North Carolina State University, PhD (Genetics) 70

Professional Experience:
Univ. Texas, Postdoc (70-72, Zoology); Univ. Michigan, Asst. to Full Prof. (72-89, Human Genetics), Prof. (85-89, Biology); Rutgers Univ., Prof. II (89-96, Marine & Coastal Sciences), Assoc. Director (89-96, Center Theoretical & Applied Genet.), Assoc. Dean – Sciences (93-95, Graduate School), Acting Assoc. Provost - Sciences (93-95, Provost’s Office), Prof. II (96-pres, Ecology, Evolution & Natural Resources, Chair 97-01)

Concurrent Positions:    
Committee – Quantitative Genetics & Common Diseases, NIH 78; Assoc Editor – Theoretical  Population Biology 79-82; Population Biology & Physiological Ecology Panel, NSF 80-83; Study Section – Mammalian Genetics, NIH 81; University of Texas, Visiting Professor of Demographic and Population Genetics 84-85; Associate Editor - Evolution 87-89; Editorial Board – International Journal of Quantitative Anthropology 88-91; Ad Hoc Committee – Endangered Amphibians, NRC 90; Associate Editor – Journal of Heredity 90-92; Human Genome Diversity Project, NIH/NSF/DoE 92; Council – American Genetic Association 94-96; Workshop – Emerging Relevance Evolutionary Biology to Applied Problems & Opportunities, Sloan/NSF94; DNA Subcommittee, New York State Forensic Commission 95-98; US National Committee – IUBS 95-98; Visiting Scholar - Australian Natl. University 96-97; Population Biology Panel – Dissertation Grants, NSF 98-99; Advisory Committee – Columbia Earth Institute 98; Scientific Advisory Board, FISHTEC 98-01; Doctoral Awards Panel, NRC/ Howard Hughes 98; Steering Committee - NJ Higher Ed. Partnership for Sustainability, Vice Pres. 00-01, Strategic Planning Committee 03-04, Review Board – Molecular Ecology 00-03; Committee on Atlantic Salmon in Maine, NRC 00-04; Advisory Board of Ecol. & Evol. Biol., Columbia University 01-pres; Advisory Board, The Land Institute 01-pres; Special Study Section, NIGMS 03; Spanish Foreign University Professor & Researcher – INIA (Spain) 04.

Biometrics & population theory, spanning the fields of evolution and ecology

Photo of Hutcheson Forest